I think I prefer the italic. It works a tiny bit better. In both the italic and the Roman the 'h' keeps jumping out at me (same as in the nasty 'Dwiggins Uncial thing'). He could have used a 'h' with an ascender. Not many words in English with 'h' and 'l' next to each other. It wouldn't have messed up his plans too much.
The 'f' and the 'g' are wonderful.
In terms of readability, though, I don't know. But then I'm not a very logical person and I don't like tomatoes.
M.